Where Photosynthesis Occurs

Photosynthesis
Plant photosynthesis occurs in leaves and green stems within specialized cell structures called chloroplasts. One plant leaf is composed of tens of thousands of cells, and each cell contains 40 to 50 chloroplasts. The chloroplast, an oval-shaped structure, is divided by membranes into numerous disk-shaped compartments. These disklike compartments, called thylakoids, are arranged vertically in the chloroplast like a stack of plates or pancakes. A stack of thylakoids is called a granum (plural, grana); the grana lie suspended in a fluid known as stroma.

Embedded in the membranes of the thylakoids are hundreds of molecules of chlorophyll, a light-trapping pigment required for photosynthesis. Additional light-trapping pigments, enzymes (organic substances that speed up chemical reactions), and other molecules needed for photosynthesis are also located within the thylakoid membranes. The pigments and enzymes are arranged in two types of units, Photosystem I and Photosystem II. Because a chloroplast may have dozens of thylakoids, and each thylakoid may contain thousands of photosystems, each chloroplast will contain millions of pigment molecules.

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Photosynthesis

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Photosynthesis, process by which green plants and certain other organisms use the energy of light to convert carbon dioxide and water into the simple sugar glucose. In so doing, photosynthesis provides the basic energy source for virtually all organisms. An extremely important byproduct of photosynthesis is oxygen, on which most organisms depend.

Photosynthesis occurs in green plants, seaweeds, algae, and certain bacteria. These organisms are veritable sugar factories, producing millions of new glucose molecules per second. Plants use much of this glucose, a carbohydrate, as an energy source to build leaves, flowers, fruits, and seeds. They also convert glucose to cellulose, the structural material used in their cell walls. Most plants produce more glucose than they use, however, and they store it in the form of starch and other carbohydrates in roots, stems, and leaves. The plants can then draw on these reserves for extra energy or building materials. Each year, photosynthesizing organisms produce about 170 billion metric tons of extra carbohydrates, about 30 metric tons for every person on earth.

Photosynthesis has far-reaching implications. Like plants, humans and other animals depend on glucose as an energy source, but they are unable to produce it on their own and must rely ultimately on the glucose produced by plants. Moreover, the oxygen humans and other animals breathe is the oxygen released during photosynthesis. Humans are also dependent on ancient products of photosynthesis, known as fossil fuels, for supplying most of our modern industrial energy. These fossil fuels, including natural gas, coal, and petroleum, are composed of a complex mix of hydrocarbons, the remains of organisms that relied on photosynthesis millions of years ago. Thus, virtually all life on earth, directly or indirectly, depends on photosynthesis as a source of food, energy, and oxygen, making it one of the most important biochemical processes known.

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Lichen

The lichen seen here growing on a tree is one of the fruticose lichens. It is made up of a layer of algal cells, a middle layer of fungal hyphae, and an outer layer of fungal tissue.
Lichen, living partnership of a fungus and an alga. The fungus component is called the mycobiont and is composed of intertwined, threadlike fibers called hyphae that are tightly packed into a tissuelike sheet. The fungus uses these hyphae to absorb food from its surroundings. The algal component, called the photobiont, makes its own food through photosynthesis and grows as a mass of green cells dispersed among the fungal hyphae. Lichens survive in a wide variety of environments, either forming small, circular crusts or leaflike structures attached to bark, rocks, or soil, or as hairlike structures hanging from tree branches.
A lichen is actually a combination of two separate organisms: an alga and a fungus. Most lichens are three-layered organisms, with an algal layer sandwiched between two layers of fungus. The alga produces the food for the lichen through photosynthesis, while the fungus absorbs water and other nutrients. Neither the fungus nor the alga can live independent of the other.
The relationship between the fungus and the alga in a lichen is an example of mutualism, in which both partners benefit from the partnership (see Symbiosis). This relationship enables each to tolerate harsh conditions where neither could survive alone. In this partnership, the fungus furnishes the alga with water, prevents overexposure to sunlight, and provides simple mineral nutrients, while the photosynthesizing alga supplies food to the fungus even if no other organic material is available. In dry, barren areas where plants have a hard time growing, such as polar tundra, deserts, rocky outcrops, or high mountains, lichens are the primary photosynthesizers. Some remarkable species even grow inside porous rocks, just below the surface where some light can still reach the algal cells.

Their ability to grow in severe conditions often makes lichens pioneers in plant succession, the process in which plants colonize bare rock or soil. Lichens release acids that break down inhospitable rock, permitting soil-trapping mosses and grasses to grab hold. In areas where soil gradually accumulates, such as a forest floor, the pioneering lichens are eventually replaced by plants and trees, although other lichens may grow on these plants and trees.

Although lichens have been used in folk medicine as purported cures for many ills, from headaches and toothaches to tuberculosis, diabetes, and asthma, their use in modern medicine is recent. The discovery in the 1940s that some fungi produce potent antibiotics stimulated an extensive screening of fungi and lichens. Since then, lichen extracts have found limited use in Europe, where lichen antibiotics have been used to treat tuberculosis and some skin diseases.


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Planting and Caring for Trees

PLANTING AND CARING FOR TREES
Whether trees are being planted for reforestation, ornament, shade, or fruit, the first step is selecting the species to be grown. The choice depends on such factors as the characteristics of the soil, the location of the site, and drainage. For example, sycamore and cottonwood trees will not grow on dry exposed slopes or ridges, or in fields with a thin topsoil over a heavy compact subsoil. Walnut trees will not grow in swampy places, and jack pines grow especially well on loose sandy soils with good drainage. A good rule of thumb is to plant native trees—trees that have demonstrated their ability to thrive in the local environment without harming other local species.

The hole for a seedling should be deep enough to hold the fully expanded root system of the seedling. Larger plants should be placed in a hole 60 cm (2 ft) deep with a diameter 60 cm greater than that of the ball of the roots. In poor soil the hole should be 1.8 m (6 ft) wide and 60 cm deep for a 2.5-m (8-ft) tree, and proportionately wider for taller trees.

After the tree is placed in the hole, the soil should be firmly pressed around the roots, and the ground should be thoroughly soaked with water. Mixing bone meal or well-rotted manure into the soil will help the tree become established quickly. Most deciduous trees should be planted in the fall when they are not growing, but evergreens are usually planted in the spring, at the beginning or middle of their period of vigorous growth.

A Watering

After planting, the soil around a tree should be kept moist, but not soaked. If artificial watering is not practical, a layer of mulch 7 to 15 cm (3 to 6 in) deep should be placed around the tree to conserve moisture and to discourage the growth of weeds. Because a transplanted tree does not adequately absorb water through its damaged roots, it is important to prevent water loss from the plant by pruning top limbs to limit transpiration.

B Fertilizing

In good soils it is less important to fertilize than in poor soils. However, all trees grow better and faster and are less likely to become diseased if fertilizer is supplied in the proper amounts. This may be done most easily by placing a large handful of fertilizer in holes made by a crowbar at the edge of the spread of the tree’s limbs. The holes should be about 60 cm (2 ft) deep and about 5 cm (2 in) in diameter, and they should be spaced about 90 cm (3 ft) apart. After the fertilizer has been introduced, the holes should be filled in with soil. Organic fertilizers such as manure and mulch are preferable to chemicals that may replace or destroy natural organisms in the soil.

C Pruning

Pruning of ornamental trees maintains the form of the tree, removes weak or sickly branches, and rejuvenates old or unhealthy plants. If performed during a period of vigorous growth, pruning often also results in an increased production of flowers. In pruning, cuts are made just above the buds that point in the direction branches are desired. When large branches are removed, the cut should be made close to the trunk, and then covered for a time.


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Major Parts of a Tree

The major parts of a tree
The major parts of a tree are its roots, trunk, leaves, flowers, and seeds. These components play vital roles in a tree’s growth, development, and reproduction.

A. Roots

Trees are held in place by anchoring organs called roots. In addition to anchoring the tree, roots also absorb water and minerals through tiny structures called root hairs. From the roots the water and mineral nutrients are carried upward through the wood cells to the leaves. Although the internal structure of most kinds of roots is similar, there are often external differences. Pines, for example, have a strongly developed taproot, or main root, in addition to branching side roots. In maples, on the other hand, there is little or no central taproot, and the other roots are produced in great numbers near the surface of the soil.

Roots grow constantly, and at the growing tip of each root is a region called the meristem, which is composed of special rapidly dividing cells. Just behind the meristem the cells become elongated, and farther from the tip the cells become differentiated into various kinds of plant tissue. In rapidly growing roots the root tip is covered by a root cap, a protective coat of loose cells that are constantly being rubbed off and replaced as the root grows.

B. Trunk

Bark is the outer protective covering of tree trunks. Because bark varies so widely in color, texture, and thickness, its characteristics provide one of the most important means of identifying species of trees. Most of the total thickness of bark consists of outer bark, which is made up of dead cells. Outer bark may be very thick, as in the cork oak, or quite thin, as in young birches and maples. Openings in the outer bark allow the movement of carbon dioxide and oxygen to and from the inner tissues.

The inner bark layer, called the phloem, consists of a thin layer of living cells. These cells act together to transport food in the form of sugars, which are made in the tree’s leaves, through the trunk and stems to other parts of the tree. Phloem cells have thin walls, and their living contents are so interconnected that the sugar solutions can pass easily and rapidly from one end of the plant to the other. As old layers of outer bark are sloughed off, new ones are constantly being added from the inside, where new phloem is always being created.

Most of a tree trunk is occupied by the wood, or xylem layer, which consists almost entirely of dead cells. The living xylem cells, however, act as the tree’s plumbing system by transporting water and dissolved food through the trunk and stems. A layer of cells called the cambium separates the living xylem cells from the phloem. As the tree grows and develops, the cambium forms new phloem and xylem cells. The layers of xylem cells form rings; these rings can be counted to determine the age of the tree in areas with distinct growing seasons.

C. Leaves

In trees, as in other green plants, the principal function of the leaves is the manufacture of sugars by the process of photosynthesis. In this process, sugars are formed when carbon dioxide (from the air) and water (from the leaf cells) are combined in the presence of light and the green pigment chlorophyll. Oxygen is produced as a byproduct. Some of the newly formed sugar is used by the leaf cells for energy, but most is carried to other parts of the tree to provide energy for growth and development in those areas.

The leaves are also the chief organs involved in the loss of water from the plant, called transpiration. Many of the tree’s tissues cannot function without a constant supply of water, and water is necessary to prevent overheating or wilting of the leaves. Transpiration is responsible for the movement of water from the roots of the tree up to the top. As water is lost through the leaves, water that enters the roots is pulled upward through the xylem tissue to replace the lost moisture, ensuring a constant circulation of water through the tissues of the tree.

D. Flowers

All angiosperms bear flowers, the trees’ reproductive structures. In some trees, such as dogwoods, cherries, and some magnolias, the flowers are large and colorful. Oaks, willows, and other temperate forest trees, on the other hand, often bear small, pale, and inconspicuous flowers.

In maples and many other trees the male and female reproductive parts are carried in separate flowers on the same tree. This arrangement is known as monoecism, and such trees are called monoecious. In oaks, for example, the male pollen-producing flowers are borne in long hanging tassels, and the short-stalked or stalkless female flowers are located on the twigs. In some trees, such as the hollies and willows, the male and female flowers are borne on separate trees. This is known as dioecism, and these trees are called dioecious.

E. Seeds

Seeds, the ripened ovules of the plant that are capable of germination, are the product of fertilized flowers and are distributed in various ways. In pines, for example, each seed is surrounded by a winglike structure. As the winged seed falls from the cone, it floats down to the ground, riding air currents. Oak seeds are enclosed in acorns, which are either planted by squirrels or merely fall to the ground near the parent tree. Willow trees produce thin-walled, flask-shaped fruits that burst open, releasing the seeds. Each seed has a tuft of downy fibers, which enables it to be picked up by air currents and travel for considerable distances. Seeds of other tree species are dispersed by water, mammals, birds, and ants.

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